Juhendajad: vanemteadur Ene Ustav (Tartu Ülikool), professor Mart Ustav (Tartu Ülikool)
Oponent: professor Karl Munger, PhD, Tufts University, Sackler School of Graduate Biomedical Sciences, Department of Developmental, Molecular and Chemical Biology, 150 Harrison Avenue, Jaharis 607, Boston MA 02111 USA.